SGI
The article sez He came out of biochemistry and genomics, where compute‑hungry (Basic Local Alignment Search Tool) (BLAST) jobs were chewing through early SGI systems.
Perhaps I'm misunderstanding the article but if I am undertanding it, the time frame is 2003-ish.
At this point you might call them "late SGI systems."
Silicon Graphics had been around for a couple of decades at that point, formed in late 1981,1
I probably first encountered them at SIGGRAPH a year or so after that.2
Really, SGI was about to become the late SGI, since it went bust in 2009.
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1 Wikipedia: Silicon Graphics
2 I remember being on a plane flying out of San Jose, California, headed to the conference and was seated across from a bunch of SGI salespeople who were loudly lamenting that SIGGRAPH was too "academically oriented" to sell much. Probably true since this was before computer graphics really broke through into the gaming, advertising, and Hollywood markets, so a majority of the attendees were broke graduate students and adjunct professors with zero purchasing power.